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 The University of North Carolina

There is growing scientific and public awareness of the widespread depletion of marine habitat-forming species, such as mangroves, seagrasses, oysters, and corals [e.g., 1,2,3]. This loss inevitably leads to the decline of the plants and animals that live in the biogenic structures created by such foundation species, and contributes to the overall degradation of marine ecosystems [4]. For example, the reduction of coral cover on tropical coral reefs directly and rapidly causes a decline in the abundance and diversity of reef fish through the loss of structural heterogeneity [5,6].